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ammianus says
11 December 1941,when Germany declared war on USA.
As Germany was already at war with Russia (from 22 June 1941, when they invaded USSR) he German declaration of war on USA made USA and USSR de facto allies.
There was no document or treaty or alliance that formally produced the alliance; Britain, USA, and USSR were all at war with Germany so cooperated as Allies to defeat Germany.
NoPlate says
The US was supplying the UK with as much war material as they could during 1941. They were also providing protection against U-boats part way across the Atlantic. This was virtually an undeclared war with Germany. US warships were sunk in this "conflict", as were U-boats.
In June 1941 (six months before Pearl Harbor and the US entrance into the war) the USSR was invaded by Germany. She was instantly in a very bad way. Britain offered help almost instantly. The USSR (again almost instantly) started to buy war goods from the US through a Soviet buying organization already in the US. This relationship between the three countries was put into writing in the Atlantic CHarter in August, 1941. The following month Lend-Lease – a plan to "lend" war supplies against future payment – was extended to the USSR in a formal manner. Lend-Lease had been going on with the UK for over a year.
In other words, the alliance started when the first bullet flew against the USSR. It was put into writing two months later.
ratcord says
There is was no official alliance as far as i know, hence the strained relations, until recently
Since they both found camps in 2008,belonging to each other on the dark side of the moon, both of them have been quiet and actually quite cordial towards each other. ( both wanted to keep that information quiet)